A somewhat modernized version is to say: it's not nature, nor nurture, it's stigmergy [1]. Or another way: the organism is larger than the collection of the cells [2] [3].
[1] "mechanism of indirect coordination, through the environment, between agents or actions. The principle is that the trace left in the environment by an individual action stimulates the performance of a succeeding action by the same or different agent", https://en.wikipedia.org/wiki/Stigmergy
[3] Michael Levin and Wayne Frasch (2023) From molecular physiology to anatomical form, https://www.youtube.com/watch?v=3l8I4gJ6D0w, at 27:32 the hybrid agent frog cells + scientists + AI exploring morphospace.
[1] "mechanism of indirect coordination, through the environment, between agents or actions. The principle is that the trace left in the environment by an individual action stimulates the performance of a succeeding action by the same or different agent", https://en.wikipedia.org/wiki/Stigmergy
[2] 2023, Michael Levin, Bioelectric networks. The cognitive glue enabling evolutionary scaling from physiology to mind, https://link.springer.com/article/10.1007/s10071-023-01780-3
[3] Michael Levin and Wayne Frasch (2023) From molecular physiology to anatomical form, https://www.youtube.com/watch?v=3l8I4gJ6D0w, at 27:32 the hybrid agent frog cells + scientists + AI exploring morphospace.