They got cross-signed by IdenTrust, which as I understand it were/are primarily selling EV certificates to financial institutions (which Let's Encrypt doesn't really compete with), rather than DV certificates (like most CAs out there do, including LE).
These days they are trusted directly by most browsers and OSes as the sibling comments mention, but the IdenTrust cross-signature was vital for bootstrapping and is still used for some older systems.